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Alpine and Subalpine Ecology Across Elevation and Climate Gradients

Explores how plant communities, soil conditions, and ecosystem function shift across elevational gradients in subalpine and alpine environments as climate change alters snowpack, temperature, and species composition.

J. HarteJ. S. LynnJohannes H. C. Cornelissenelevational gradientsspace usespecies diversityNEON AOP Survey of Upper East River CO Watersheds:Range size and local abundance data for angiospermData from: 'Abiotic influences on continuous conifThe Impacts of Large Houses: A Recommendation to Lquadrat sampling method5TE soil sensor monitoringSpecific Leaf Area (SLA)Winters are changing: snow effects on Arctic and aThe structure and function of subalpine ecosystemsEcosystem responses to warming-induced plant speciAmerican Farmland TrustSonoran InstituteGunnison County Assessor's Office

Knowledge Graph (870 nodes, 3951 connections)

Research Primer

Background

Mountain ecosystems compress dramatic environmental change into short vertical distances. In the Gunnison Basin of Colorado, a hike of a few hundred meters up a ridge can cross differences in temperature, snowpack, growing season length, and soil development that would otherwise require hundreds of kilometers of latitudinal travel. Researchers exploit these elevational gradients — spatial transects along which temperature, snowmelt timing, and soil properties all change predictably — as natural experiments for understanding how alpine and subalpine communities are assembled and how they might respond to a warming climate. Because the high-elevation meadows, forests, and tundra around the Rocky Mountain Biological Laboratory (RMBL) are cold-limited and seasonally snow-covered, even small shifts in temperature or snowmelt date can reshuffle which species grow where, when they flower, and how they interact.

Several key concepts recur throughout this body of work. Community composition — the identity and relative abundance of species in a place — is shaped not only by climate but by biological interactions, including those with arbuscular mycorrhizal fungi and other fungal endophytes that live inside plant roots and leaves and influence nutrient uptake and stress tolerance. Plant functional groups (for example, forbs, grasses, and shrubs) group species by shared ecological strategies, and shifts among these groups — especially shrub encroachment, where woody plants expand into meadows — are important signals of ecosystem change. Phenology, particularly flowering time, is tightly tied to snowmelt date, making it a sensitive indicator of climate warming. At larger scales, ecologists use the species-area relationship (S = cA^z, where species richness scales with sampled area) and related tools from the maximum entropy theory of ecology to predict biodiversity patterns from small plots up to whole landscapes.

These ideas matter for the Gunnison Basin because its subalpine meadows, aspen groves, and alpine ridges provide forage for livestock and wildlife, store substantial soil carbon, support pollinators, and underpin the region's tourism and water supply. Understanding how warming, changing snow regimes, and altered species interactions propagate through these systems is essential for land managers anticipating shrub expansion, shifts in meadow productivity, and changes in wildflower displays.

Foundational work

Much of the modern experimental tradition at RMBL traces to a pioneering meadow-warming experiment begun in the early 1990s, in which overhead infrared radiators added a constant 15 W/m² of heat to subalpine meadow plots (Harte et al., 1995). This manipulation advanced snowmelt by roughly a week, raised summer soil temperatures by up to 3°C, and reduced soil moisture by as much as 25%, establishing a template for climate-warming experiments worldwide and providing a conceptual framework that others would adopt for interpreting ecosystem warming responses (Shaver et al., 2000). Subsequent work at the same site showed that warming advanced flowering by about 14 days, with early-flowering species such as Claytonia lanceolata most strongly affected, and demonstrated that snowmelt date was the single most important microclimatic driver of phenological change (Dunne et al., 2003).

Parallel foundational research expanded these insights to broader scales. Cross-biome syntheses showed that ecosystems converge to a common maximum rain-use efficiency under the driest conditions (Huxman et al., 2004), and mountain studies revealed that bacterial and plant diversity follow strikingly different elevational patterns: bacterial richness declined monotonically with elevation while plant richness peaked at mid-elevations (Bryant et al., 2008). On the theoretical side, the maximum entropy theory of ecology provided a unifying framework for predicting species-area curves, abundance distributions, and energetic patterns from a small number of community-level constraints (Harte et al., 2009; Harte & Newman, 2014) (Harte & Newman, 2014).

Key findings

Decades of experimental warming at RMBL and comparable sites have produced a consistent picture: warming advances snowmelt, dries soils, and reorganizes plant communities toward woodier, more conservative vegetation. A 23-year synthesis found that experimental warming advanced snowmelt by an average of 7.7 days and accelerated snowmelt by an additional half-day per year, linking these microclimate changes to shrub encroachment and transient soil carbon loss (convergent ecosystem responses, 2015). Drought- and warming-induced shifts from forbs to shrubs drove substantial soil carbon losses through declines in community productivity (shifts in plant dominance, 2006), and a 10-year heating experiment reduced soil organic matter by roughly 200 g C/m² (plant community composition mediates soil carbon, 2002). At the biome scale, community-level plant height has increased with warming across tundra sites, although other functional traits have lagged behind expected rates of change (Bjorkman et al., 2018), and tundra vegetation surveys confirm that these plot-scale changes are linked to recent summer warming (Elmendorf et al., 2012).

Belowground, the story is equally striking. Warming drying tends to reduce soil moisture across experiments, but the temperature sensitivity of soil respiration itself has remained largely unchanged, with a universal decline in sensitivity above 25°C (Carey et al., 2016). Global syntheses show that warming-induced soil carbon losses depend strongly on the size of initial carbon stocks, with high-latitude and high-elevation systems especially vulnerable (Crowther et al., 2016), though models and experiments still disagree on direction and magnitude (Sulman et al., 2018). Plant-fungal symbioses are central to these responses: fungal symbionts alter plant responses to nearly every global change factor except elevated CO₂, with arbuscular mycorrhizal and ectomycorrhizal fungi generally buffering plants most strongly (Kivlin et al., 2013). At RMBL, warming has been shown to cause local extinction of sensitive forbs such as Androsace septentrionalis by depressing fecundity and survival (climate warming drives local extinction, 2018), while species that persist often do so through compensatory growth by deeper-rooted neighbors (compensatory responses, 2007).

Warming also reshapes biotic interactions beyond symbioses. Herbivore attack on subalpine grasses increased by 13% and damage by 29% under experimental warming, with effects contingent on fine-scale weather (Lynn et al., 2023). Ant communities have shifted upslope over 13 years of natural climate change, with modern high-elevation assemblages resembling historical low-elevation ones (changes in ant community composition, 2014). Meanwhile, theoretical work has expanded the species-area relationship into a universal curve that lets ecologists upscale biodiversity estimates from small plots to entire landscapes (Harte et al., 2009) and predict regional arthropod richness from a handful of tiny census plots (inferring regional-scale species diversity, 2015).

Current frontier

Early work in the 1990s and 2000s focused on documenting whether and how warming changes montane plant communities. Research since 2020 has shifted toward mechanisms — especially the coupling between plants and their microbial partners — and toward whole-community responses along elevational gradients. A 2026 synthesis from long-running RMBL experiments shows that warming caused a 150% increase in shrub cover and roughly 28% declines in forbs and grasses, while mycorrhizal and septate root colonization both dropped 17–20%, effectively decoupling above- and belowground communities (Souza et al., 2026). A multi-site study spanning 2,000 km and 12°C of mean annual temperature found that warming reduced septate fungal colonization by 90% in leaves and 35% in roots, and weakened the coordination between fungal communities and plant metabolism (Edwards et al., 2025). A whole-community transplant experiment across a 400-meter elevation gradient revealed that soil bacteria and archaea respond fastest to warming, followed by fungi and then plants, hinting at emerging mismatches between partners that historically co-occurred (Seltzer, 2025).

Other recent work is expanding the taxonomic and ecological breadth of the neighborhood. Studies are examining the contributions of rare species to phylogenetic diversity in subalpine meadows (Veldhuisen et al., 2025), the role of hemiparasitic Castilleja in boosting plant species richness (Hastings, 2024; Lang, 2023) (Lang, 2023), and the distinct biodiversity signature of beaver-engineered wetlands (McDonough, 2024). At the same time, ecologists are finding that static macroecological theory increasingly fails in stressed alpine plant communities, with species-area and abundance patterns drifting from maximum entropy predictions over just six years (shifting macroecological patterns, 2021). Methodological advances — DNA metabarcoding of fungal communities, drone-based vegetation mapping, and reactive transport models linking biogeochemistry to hydrology (tracking diverse minerals, 2019) — are opening new windows on these changes.

Open questions

Several major uncertainties remain. How will decoupling of plant-fungal symbioses cascade through community assembly, nutrient cycling, and carbon storage over coming decades? Can process-based soil carbon models be reconciled with the wide range of observed responses in warming experiments (Sulman et al., 2018), and when do montane meadows switch from carbon sinks to carbon sources? Why do static macroecological theories increasingly fail in stressed alpine communities, and can these deviations themselves become useful early-warning indicators of ecosystem reorganization? How will shrub encroachment, shifting pollinator assemblages, and upslope movement of species interact to produce novel communities in the Gunnison Basin? Addressing these questions will require continued long-term experiments, integration of above- and belowground monitoring, and theoretical frameworks flexible enough to handle non-stationary, rapidly changing systems.

References

Bjorkman, A.D. et al. (2018). Plant functional trait change across a warming tundra biome. Nature.

Bryant, J.A., Lamanna, C., Morlon, H., Kerkhoff, A.J., Enquist, B.J., Green, J.L. (2008). Microbes on mountainsides: contrasting elevational patterns of bacterial and plant diversity. PNAS.

Carey, J.C. et al. (2016). Temperature response of soil respiration largely unaltered with experimental warming. PNAS.

Changes in ant community composition caused by 20 years of experimental warming vs. 13 years of natural climate shift (2014).

Climate Warming Drives Local Extinction: Evidence from Observation and Experimentation (2018).

Compensatory responses to loss of warming-sensitive plant species (2007).

Convergent ecosystem responses to 23-year ambient and manipulated warming (2015).

Crowther, T.W. et al. (2016). Quantifying global soil carbon losses in response to warming. Nature.

Dunne, J.A., Harte, J., Taylor, K.J. (2003). Subalpine meadow flowering phenology responses to climate change: integrating experimental and gradient methods. Ecological Monographs.

Edwards, K.F. et al. (2025). Warming disrupts plant-fungal endophyte symbiosis more severely in leaves than roots. Global Change Biology.

Elmendorf, S.C. et al. (2012). Plot-scale evidence of tundra vegetation change and links to recent summer warming. Nature Climate Change.

Harte, J., Newman, E.A. (2014). Maximum information entropy: a foundation for ecological theory. Trends in Ecology and Evolution.

Harte, J., Smith, A.B., Storch, D. (2009). Biodiversity scales from plots to biomes with a universal species-area curve. Ecology Letters.

Harte, J., Torn, M.S., Chang, F.-R., Feifarek, B., Kinzig, A.P., Shaw, R., Shen, K. (1995). Global warming and soil microclimate: results from a meadow-warming experiment. Ecological Applications.

Hastings, E. (2024). Getting to the Root of It: Effects of Castilleja Root Hemiparasitism on Plant Community Structure and Function.

Huxman, T.E. et al. (2004). Convergence across biomes to a common rain-use efficiency. Nature.

Inferring Regional-Scale Species Diversity from Small-Plot Censuses (2015).

Kivlin, S.N., Emery, S.M., Rudgers, J.A. (2013). Fungal symbionts alter plant responses to global change. American Journal of Botany.

Lang, A. (2023). Host specificity of hemiparasitic Castilleja and its influence on plant community diversity.

Lynn, J.S. et al. (2023). Herbivory damage but not plant disease under experimental warming is dependent on weather for three subalpine grass species. Journal of Animal Ecology.

McDonough, L. (2024). Environmental Variation and Vegetative Composition in Beaver Engineered Ecosystems.

Plant community composition mediates both large transient decline and predicted long-term recovery of soil carbon under climate warming (2002).

Seltzer, C. (2025). The impacts of environmental change on plant and microbial communities: A turf transplant experiment in the Colorado Rocky Mountains.

Shaver, G.R. et al. (2000). Global warming and terrestrial ecosystems: a conceptual framework for analysis. BioScience.

Shifting macroecological patterns and static theory failure in a stressed alpine plant community (2021).

Shifts in plant dominance control short and long-term carbon-cycle responses to widespread drought (2006).

Souza, L. et al. (2026). Experimental warming decouples plant-fungal symbiont interactions and leads to a more conservative ecosystem. PNAS.

Sulman, B.N. et al. (2018). Multiple models and experiments underscore large uncertainty in soil carbon dynamics. Biogeochemistry.

Tracking diverse minerals, hungry organisms, and dangerous contaminants using reactive transport models (2019).

Veldhuisen, L. et al. (2025). Rare species do not disproportionately contribute to phylogenetic diversity in a subalpine plant community. American Journal of Botany.

Concept (78) →

elevational gradients

Spatial gradients in elevation that capture variation in temperature, soil age and type, disturbance regimes, and land-use histories providing importa...

frameworkgeneral ecology392 papers

space use

Way in which individuals distribute themselves across landscape, including movement patterns and spatial organization like home range size and overlap

measurementlandscape310 papers

species diversity

A measure of community diversity that accounts for both species richness and relative abundance, calculated using the Shannon-Weiner index

metriccommunity ecology240 papers

nutritional niche partitioning

The concept that co-occurring species occupy distinct positions in multidimensional nutrient space to minimize competition and facilitate coexistence

theorycommunity ecology94 papers

keystone species

A species that has a disproportionately large effect on its environment relative to its abundance and whose presence/absence significantly affects eco...

theorycommunity ecology72 papers

arbuscular mycorrhizal fungi

Fungal symbionts that form specialized structures (arbuscules, vesicles) within plant root cells and are important for nutrient exchange

measurementcommunity ecology69 papers

Species-area relationship

The positive relationship between species richness and the surveyed area, described by the power-law S=cA^z where c is the intercept and z measures th...

phenomenoncommunity ecology61 papers

permafrost

phenomenonclimate60 papers

spatial clustering

Alpine plants growing in clumps with multiple species interacting in close proximity, affecting microhabitats and species interactions

phenomenoncommunity ecology57 papers

shrub encroachment

Shift toward woody plant encroachment into nonwoody meadows and grasslands

phenomenoncommunity ecology47 papers
Show 68 more concepts

trait filtering

Environmental processes that filter trait values admitted by changing abiotic and biotic conditions, maintaining trait distributions through phenotypi...

processpopulation ecology47 papers

breeding ecology

The study of reproductive behaviors, nesting habitat selection, and breeding success in bird populations

processpopulation ecology42 papers

chlorophyll content

SPAD measurement of leaf chlorophyll concentration indicating photosynthetic capacity

measurementgeneral ecology38 papers

ecosystem engineering

The modification of environments by organisms in ways that influence resource availability for other species, exemplified by beaver dam construction c...

processcommunity ecology37 papers

maximum entropy theory of ecology

Theory that predicts patterns of distribution, abundance, and energetics using instantaneous values of community state variables including total area,...

theorycommunity ecology35 papers

climate-ecosystem feedback

Ecosystem responses to climate change that exert positive or negative feedbacks on climate, mediated by slow-moving factors such as shifts in vegetati...

processclimate33 papers

dark septate endophytes

A group of fungal endophytes characterized by dark-pigmented, septate hyphae that commonly colonize plant roots and can increase under environmental s...

measurementcommunity ecology32 papers

root weight fraction

The ratio of dry root weight to total dry plant weight, used as an indicator of soil nutrient availability with higher values indicating nutrient-poor...

metricmethodological31 papers

community composition

The identity and relative abundance of species within an ecological community

measurementcommunity ecology30 papers

secondary cavity nesting

Bird species that utilize nest cavities created by other species rather than excavating their own

processcommunity ecology24 papers

pollinator visual space

Mathematical models that represent how different pollinators perceive color based on their photoreceptor sensitivities and opponent color processing

frameworkcommunity ecology22 papers

plant functional groups

Classification of species with similar characteristics into plant functional groups or plant functional types to reduce complexity in ecological commu...

frameworkcommunity ecology20 papers

territoriality

Territorial behavior including alarm and warning calls to protect midden from squirrels and other mammals, with territories ranging from 0.5 to 1 hect...

phenomenonpopulation ecology19 papers

indicator species

Taxa that are significantly associated with particular habitat conditions and can be used to assess ecosystem state or restoration success

measurementcommunity ecology19 papers

climatic niche

The range of climate conditions that a species can tolerate, estimated from ecological niche models

conceptgeneral ecology17 papers

metabolic rate distribution over individuals

Distribution of metabolic rates over individuals in a community, obtained by summing over abundance

measurementcommunity ecology15 papers

hemiparasitism

Parasitic plants that retain absorptive roots and are capable of carbon fixation through photosynthesis but still need to draw some of their resources...

processcommunity ecology15 papers

host specificity

The degree to which parasites show preference or restriction to particular host species rather than being generalist

phenomenoncommunity ecology15 papers

plant acclimation

Physiological shifts that allow plants to adjust their rates of photosynthesis and stomatal conductance to compensate for changes in temperature

processgeneral ecology15 papers

invasive species establishment

The process by which non-native species become established and spread in new environments

processcommunity ecology14 papers

maximum entropy inference

Statistical inference procedure that selects flattest probability distributions compatible with constraints imposed by prior knowledge

frameworkmethodological13 papers

phylogenetic diversity

A measure of the evolutionary history represented in an ecological community, calculated as the sum of branch lengths connecting species in a phylogen...

measurementcommunity ecology12 papers

litter quality

Chemical composition of plant litter that determines its decomposability, typically characterized by C:N ratios and lignin content

measurementgeneral ecology11 papers

ecotonal zones

Areas of transition between adjacent, distinct ecosystems with unique species assemblages at the fringes of mature stands

phenomenonlandscape11 papers

host-microbiome interactions

The relationship between a host organism and its associated microbial community, including costs and benefits to host fitness

frameworkcommunity ecology11 papers

behavioral ontogeny

processgeneral ecology11 papers

information entropy

A quantitative measure of uncertainty about an outcome of a draw from a probability distribution

metricmethodological11 papers

species accumulation curve

Plot showing cumulative number of species discovered as a function of sampling effort, used to assess sampling completeness

measurementcommunity ecology10 papers

plant disease

Percentage of leaf area affected by pathogen symptoms including powdery mildew, rust, and other fungal diseases

measurementcommunity ecology10 papers

pioneer species

Species best suited for the environment that make up most of the vegetation in early successional stages

conceptcommunity ecology8 papers

mass ratio hypothesis

Species with greater primary production exert the main controls for the functioning of ecosystems due to greater aboveground abundance of biomass or l...

hypothesiscommunity ecology8 papers

faunal elements

Biogeographic classification of species based on their broader distributional affinities and evolutionary origins

theorybiogeography8 papers

snow mosquitoes

Univoltine species which develop from overwintered eggs in pools formed by melting snow water

phenomenongeneral ecology8 papers

competitive release

Hypothesis that parasitism of dominant plant species reduces their competitive ability, allowing less competitive species to increase in abundance

hypothesiscommunity ecology7 papers

electrolyte leakage

Loss of cellular electrolytes through compromised cell membranes, used as indicator of heat stress damage

measurementmethodological7 papers

ecological restoration

The process of managing or assisting the recovery of an ecosystem that has been degraded, damaged or destroyed as a means of sustaining ecosystem resi...

processgeneral ecology7 papers

high-temperature tolerance

Ability of plants to survive exposure to elevated temperatures, measured as lethal temperature causing 50% electrolyte leakage (LT50)

measurementgeneral ecology7 papers

herbarium collections

Preserved plant specimens that are picked in the field, pressed flat, dried and attached to archival-grade paper with relevant information recorded, t...

measurementmethodological6 papers

population decline

Reduction in population growth rates, size, or occurrence across species ranges

phenomenonpopulation ecology6 papers

energy equivalence principle

The principle asserting an inverse relationship between local population density and average metabolic rate of individuals in a species

hypothesispopulation ecology6 papers

montane ecosystem

Mountain ecosystem characterized by specific elevation, climate, and vegetation adapted to montane conditions

habitat typelandscape6 papers

soil sterilization

Experimental technique to remove soil microbial communities via autoclaving

methodologymethodological5 papers

local resource competition

Theory suggesting that offspring sex ratios might vary according to maternal condition, hence females should vary the sex of their offspring according...

theorypopulation ecology5 papers

ideal free distribution

Theory that animals distribute themselves among habitat patches in proportion to resource availability, with high-quality patches becoming occupied fi...

theorypopulation ecology5 papers

Fisher logseries distribution

A statistical distribution commonly observed in species abundance patterns, predicted by METE for undisturbed communities

model typecommunity ecology5 papers

distance-decay relationship

Decrease in compositional similarity between communities with increasing geographic distance or environmental separation

phenomenoncommunity ecology5 papers

competitive exclusion

Competition between fungal groups for limited plant photosynthate allocation

processcommunity ecology4 papers

internode length

The distance between branch whorls on conifers, used to measure annual growth

measurementmethodological4 papers

transplanting

Moving native plants from one location to establish them in a disturbed site as a revegetation technique

processlandscape4 papers

point count surveys

Standardized method for observing and counting birds at fixed locations for specified time periods

measurementmethodological4 papers

upscaling

Extrapolation of species richness estimates from small plots to larger spatial scales using theoretical scaling relationships

processmethodological4 papers

keystone species complex

Sapsucker-fungus partnership that strongly influences the avian community by providing nest sites

theorycommunity ecology3 papers

aspect effect

Differences in environmental conditions and species composition between north and south-facing slopes

processlandscape3 papers

Constant Connectance in Space model

Model developed to determine LARs based on SAR and simple properties of food webs including species richness, number of links and connectance

model typecommunity ecology3 papers

soil horizon

Distinct layers in soil profile with different physical and chemical properties

measurementbiogeochemistry3 papers

standardized effect size

A metric that compares observed values to null model expectations, calculated as (observed - expected) / standard deviation of null distribution

measurementmethodological3 papers

Gloger's rule

A relationship between plumage coloration and climate variation, predicting that darker coloration due to pigmentation occurred in warmer, more humid ...

hypothesisgeneral ecology2 papers

theory compression

processmethodological2 papers

microbial biogeography

The study of spatial distribution patterns of microorganisms and the factors controlling these patterns

frameworkcommunity ecology2 papers

Relative Interaction Intensity

Standardized measure of interaction strength ranging from -1 to 1, where positive values indicate facilitation and negative values indicate competitio...

metriccommunity ecology2 papers

nearest neighbor analysis

Statistical method comparing species composition near target individuals to background composition to identify spatial associations

methodmethodological2 papers

temperature lapse rate

The rate at which temperature decreases with elevation, typically expressed as degrees Celsius per kilometer

measurementclimate2 papers

herbarium specimen collection bias

Systematic tendency for historical plant collectors to preferentially collect rare or uncommon species while avoiding common, dominant species

phenomenonmethodological2 papers

photo-inhibition

Reduction in photosynthetic efficiency due to excess light exposure

processgeneral ecology2 papers

Dobzhansky-MacArthur hypothesis

Predicts that a species' high-elevation range limit is determined by harsh abiotic conditions whereas its low-elevation range limit is set by antagoni...

hypothesisbiogeography2 papers

secondary sexual characteristics

Traits like crown-stripe width, plumage coloration, and song that require additional energy to maintain and signal mate quality

phenomenonevolution2 papers

Simpson diversity

metriccommunity ecology2 papers

lambda

metricpopulation ecology2 papers

Protocol (64) →

quadrat sampling method

Measurement of plant species-specific cover, identification of focal species, and collection of plant biomass and functional traits including specific...

samplingstandardized161 papers

Specific Leaf Area (SLA)

Comprehensive measurement of vegetative traits including specific leaf area, trichome density, water use efficiency, and percent water content from fi...

measurementstandardized87 papers

5TE soil sensor monitoring

Continuous automated monitoring of soil temperature, moisture, and gas concentrations at multiple depths using sensor arrays. Provides high-resolution...

measurementstandardized87 papers

DNA metabarcoding

Surface sterilization of plant tissues followed by DNA extraction and high-throughput sequencing of the ITS region to characterize fungal endophyte co...

analyticalstandardized80 papers

Holshof & Swenson trait sampling protocol

Systematic monitoring of plant disease dynamics across elevation gradient to capture environmental variation effects. Involves establishment of standa...

observationalstandardized71 papers

McGonigle method (Poaceae)

Microscopic assessment of arbuscular mycorrhizal fungal colonization and other fungal colonization of plant roots using crosshair intercept method and...

analyticalstandardized63 papers

Infrared heating manipulation

Infrared heaters suspended above experimental plots to simulate warming effects on montane vegetation communities across moisture gradients. Heating p...

experimentalstandardized52 papers

HOBO sensor environmental monitoring

Continuous collection of local weather data at 5-minute intervals throughout growing seasons to capture fine-scale environmental variation affecting d...

measurementstandardized48 papers

RMBL warming experiment

29-year experimental warming using overhead infrared radiators to increase soil temperature by 2°C in subalpine grassland plots. Heat flux of 22 W/m² ...

experimentalstandardized47 papers

reflectance spectrophotometry

Standardized measurement of floral reflectance spectra using fiber optic probe positioned at fixed distance and angle from flower surface. Spectra col...

measurementstandardized40 papers
Show 54 more protocols

Point count surveys

Systematic point count surveys conducted along an elevational gradient to assess current species distributions. Each site has four sampling points wit...

observationalstandardized36 papers

Li-Cor 6400 photosynthesis measurement

A novel approach to measure light response curves for entire plant communities rather than individual plants using tent-based gas exchange measurement...

measurementstandardized33 papers

Mantel test

Calculation of multiple phylogenetic diversity metrics for plant communities using established phylogenetic trees and null model approaches to test fo...

analyticalstandardized28 papers

SPAD chlorophyll measurement

Measurement of chlorophyll fluorescence using LI-600 to assess photosynthetic efficiency by quantifying light emission from chlorophyll molecules exci...

measurementstandardized27 papers

MaxEnt SAR theory

Application of METE mathematical framework to predict species-area relationships and species-abundance distributions based on community state variable...

computationalstandardized24 papers

Alpine soil inocula collection and preparation

Collection of soil inocula from resident (high elevation) and novel (low elevation) sites across elevation gradients in mountain ecosystems, with subs...

samplingstandardized24 papers

soil incubation (Asteraceae)

Factorial greenhouse experiment testing effects of soil inoculum source and sterilization on alpine grass growth, traits, and fungal colonization. Pla...

experimentalstandardized21 papers

Beaver pond habitat characterization

Multi-parameter assessment of beaver pond physical characteristics including area, depth, flow velocity, and beaver activity indicators to control for...

measurementstandardized20 papers

Nondestructive plant size measurement for biomass prediction (Asteraceae)

Systematic measurement of plant morphological characteristics (leaf number, leaf area, stalk count, etc.) before plant extraction to develop predictiv...

measurementstandardized19 papers

Subalpine plant community energetics census

Complete census of all individual plants in gridded plots with spatial mapping, tissue sampling, and photosynthetic area measurement for metabolic rat...

samplingstandardized17 papers

Historical herbarium database analysis for biographical reconstruction

Method for extracting and analyzing historical herbarium specimen data to reconstruct the life and career patterns of past collectors, including tempo...

analytical16 papers

species-area relationship

Nested sampling design to construct species-area relationships across multiple spatial scales by subdividing plots into equal-area quadrats and averag...

analyticalstandardized15 papers

Elevational fungal transplant experiment (Poaceae)

A controlled field experiment transplanting grass species across elevational gradients with manipulated fungal soil communities to test effects of pla...

experimental15 papers

percent cover estimation

Long-term monitoring of tundra plant communities using standardized sampling within permanently marked study areas, employing various abundance quanti...

samplingstandardized14 papers

community-weighted mean

Multivariate analysis technique to quantify community responses over time. Treatment weights indicate response magnitude and species weights show indi...

analytical14 papers

destructive biomass sampling

Complete excavation of plants followed by separation into above-ground, below-ground, and reproductive components, with subsequent drying and weighing...

samplingstandardized12 papers

Elevation gradient flower spectrophotometry

Systematic collection and spectrophotometric analysis of flowers across elevation gradients to quantify color variation. Targets pollinator-visible fl...

measurement12 papers

Reciprocal transplant germination experiment

Field experiment testing seed germination of different genotypes/treatments in natural microhabitats of related species to assess niche requirements a...

experimental11 papers

Grid-based root presence-absence quantification from soil pit photographs (Plantae)

Photographs of soil pit faces are overlaid with 1×1 cm grids to systematically score presence or absence of fine and coarse roots in each grid cell, p...

measurementstandardized11 papers

gravimetric soil moisture analysis

Standard gravimetric determination of soil moisture content using oven-drying method at 100°C for 24 hours with 2mm sieving to remove rocks and organi...

laboratory11 papers

GIS polygon digitization

Using ArcGIS to digitize habitat polygons and generate random sampling points within appropriate community types.

analytical10 papers

Plant specimen collection and identification

Pressing and identification of plant voucher specimens using standard botanical keys and herbarium references.

laboratory10 papers

Potter trap

Standard bird capture protocol using Potter's traps baited with seed, followed by individual banding and morphological measurements for population mon...

sampling10 papers

CDC light trap sampling (Culicidae)

Opportunistic collection of adult and larval mosquitoes across elevational gradients using standardized collection techniques. Adults collected during...

sampling9 papers

Endophyte removal using Benomyl fungicide (Poaceae)

Seeds were treated with Benomyl fungicide at 4°C to remove endophytic fungi, creating endophyte-free plants for comparison with naturally endophyte-po...

experimental8 papers

Adult mosquito collection and identification (Culicidae)

Collection of adult mosquitoes during blood-seeking behavior and other activities to complement larval surveys and confirm species identifications.

sampling8 papers

Diurnal thermal videography of plant communities

Continuous infrared thermal imaging of vegetation plots at 5-second intervals from sunrise to sunset using elevated FLIR cameras to capture leaf tempe...

measurement8 papers

NRCS soil characterization methods (Plantae)

Random plot establishment along elevational gradients followed by soil coring and root extraction to quantify coarse and fine root biomass allocation ...

samplingstandardized8 papers

Paired plot community comparison (Orobanchaceae)

Vegetation composition and diversity are compared between paired 1m² plots with and without Castilleja presence to assess community-level effects of p...

sampling8 papers

Habitat-damage correlation analysis (Aves)

Quantitative analysis of the relationship between sapsucker damage patterns on aspen trees and proximity to willow habitat, using distance measurement...

analytical8 papers

Cavity nest site characterization (Aves)

Comprehensive measurement of nest tree characteristics and landscape context including tree size, nest height, cavity orientation, and distances to ke...

measurement7 papers

Successional stage soil sampling across landslide chronosequence

Systematic sampling of soil from different plant successional stages within a landslide area to compare soil properties across a chronosequence. Uses ...

sampling7 papers

Electrolyte leakage assay (Asteraceae)

Controlled temperature treatments from 15-65°C applied to detached leaves for 1 hour, with temperature ramped at 2°C/hour. Used to determine lethal te...

experimental7 papers

U.S. Geological Survey sediment preparation procedures

Extraction of iron and manganese from sediments using 0.5 N HCl with hydroxylamine hydrochloride in anaerobic conditions, followed by ICP-AES analysis...

analyticalstandardized6 papers

Nearest neighbor host detection analysis (Orobanchaceae)

Plant species composition within 15cm of Castilleja individuals is compared to random background vegetation to statistically identify overrepresented ...

analytical6 papers

GIS-based riparian proximity measurement (Picidae)

Measurement of distances from bird survey points to nearest riparian features using aerial photography, field measurements, and GIS analysis to quanti...

measurement6 papers

Species-area curve analysis

Systematic sampling of lichen cover and species composition along fixed 1-meter transects on rock surfaces, with digital photography and image analysi...

sampling6 papers

song playback experiment (Passeridae)

Systematic presentation of conspecific, heterospecific competitor, and control songs to territorial males followed by standardized behavioral observat...

experimental5 papers

oven-dry biomass determination

Standardized protocol for determining dry mass of plant tissues through controlled oven drying at 60°C for 48 hours followed by precision weighing to ...

laboratory5 papers

Grid-based topographic measurement

Systematic measurement of slope and aspect at grid cell centers using compass and clinometer with clipboard to minimize micro-scale surface variation ...

measurement4 papers

bird banding (Passerellidae)

Standard measurements of bird morphology including wing chord, tail length, tarsus length, and body condition assessment. Birds banded with color comb...

measurement4 papers

Riparian willow structural complexity assessment (Parulidae)

Combined vertical (Robel pole) and horizontal (line intercept) measurements to quantify willow habitat structural complexity including height diversit...

measurement4 papers

Shannon Diversity Index

Calculation of Shannon diversity index (ln type) and Simpson dominance index (1-D) from species count data using online biodiversity calculator. Stand...

analytical3 papers

Elevational gradient soil and seed sampling (Fabaceae)

Systematic collection of soils and seeds across elevational gradients spanning current and projected future species ranges to capture natural variatio...

sampling3 papers

Castilleja haustorial connection microscopy (Orobanchaceae)

Root systems around Castilleja plants are excavated, cleaned, and examined microscopically to identify haustorium-host connections for direct detectio...

analytical3 papers

LC-MS plant metabolomics (Poaceae)

Extraction of plant metabolites using methanol/acetonitrile/water solvent followed by liquid chromatography-mass spectrometry analysis. Characterizes ...

analytical3 papers

pavo visual modeling

Conversion of reflectance spectra into bee and bird visual spaces using established photoreceptor sensitivity models. Uses color hexagon model for bee...

computational3 papers

Lichen field identification

Field identification of lichen species using chemical spot tests with potassium hydroxide, morphological examination with hand lens, and comparison to...

analytical3 papers

Multi-model soil carbon simulation

Comparative simulation study using five different soil organic carbon models (DAYCENT, CORPSE, MIMICS, MEND, RESOM) to project responses to warming an...

computational3 papers

Rolatape transect gopher disturbance survey

Systematic survey of pocket gopher soil disturbances (eskers, mounds) along standardized transects using rolatape measuring wheels with perpendicular ...

sampling2 papers

Systematic grid mapping of sagebrush populations (Asteraceae)

Complete census of sagebrush individuals within systematic 16m × 16m grids divided into 1m² cells, with measurement of plant dimensions for size estim...

sampling2 papers

Ion exchange resin strips

Mixed-bed ion-exchange resin bags deployed at 5 cm soil depth to assess NO3-N and NH4-N availability. Resins are extracted with KCl and analyzed using...

measurement2 papers

IUCN Red List Area of Occupancy calculation

Calculation of species area of occupancy (AOO) using filtered GBIF occurrence records and standardized grid-based methods following IUCN guidelines fo...

analytical2 papers

Length-mass relationship conversion

Estimation of invertebrate biomass using established taxon-specific length-mass relationships and scaling to aerial measures.

analytical2 papers

Stakeholder (3)

American Farmland Trust

ngo3 docs

Sonoran Institute

academic2 docs

Gunnison County Assessor's Office

local gov2 docs

Author (97) →

Show 87 more authors

E. J. Cooper

3 works

D. Blok

2 works

L. M. Kueppers

University of California BerkeleyORCID: 0000-0002-8134-357913 works

S. R. Saleska

8 works

M. E. Loik

7 works

E. Dorrepaal

2 works

E. Newman

6 works

G. Kudo

3 works

L.A. Souza

3 works

J. A. Dunne

4 works

R. G. Bjork

2 works

R. D. Hollister

2 works

C. F. Williams

Rocky Mountain Biological LaboratoryORCID: 0000-0003-4138-308X14 works

I. S. Jonsdottir

2 works

J. M. Welker

2 works

T. A. Day

2 works

W. A. Gould

2 works

H. Marshall Worsham

Lawrence Berkeley National LaboratoryORCID: 0000-0001-7924-040X13 works

A. D. Bjorkman

3 works

M. Brush

3 works

D. L. Taylor

2 works

M. L. Fischer

3 works

K. Umemura

3 works

Kenneth D. Whitney

6 works

Brieanne Forbes

Pacific Northwest National Laboratory (PNNL)ORCID: 0000-0001-9240-96874 works

A. M. Panetta

6 works

C. L. Debban

2 works

M. H. Koski

3 works

A. Rominger

3 works

T. L. Ashman

2 works

M. Shaw

4 works

Nisreen Abo-Sido

2 works

J. L. Kortmeyer

2 works

Sabastian Escobar

3 works

M. Gray

2 works

J. Kitzes

5 works

R. M. Rizo

3 works

N. E. Krauss

2 works

A. Kinzig

4 works

Shermila B. Villanueva

2 works

C. Levy

2 works

K. A. Stinson

5 works

K. J. Taylor

7 works

R. E. Poore

2 works

C. B. Sides

2 works

T. E. White

2 works

Leah N. Veldhuisen

University of ArizonaORCID: 0000-0002-7094-01086 works

A. N. Trautwig

2 works

C. Young

2 works

T. Huxman

2 works

Y. Zhang

3 works

L. Novak

2 works

M. S. Torn

4 works

G. R. Shaver

2 works

Ross R. Conover

3 works

R. Seagrist

6 works

Samuel Canfield

3 works

E. Hoffman

3 works

K. K. Beals

2 works

W. Hendricks

2 works

L. B. Ranelli

2 works

M. J. Stansberry

2 works

M. Wilber

2 works

Katrina M Dlugosch

ORCID: 0000-0002-7302-66372 works

D. Cohen

2 works

D. Bartholomew

2 works

K. Alvarado

2 works

D. Levy

2 works

A. Adams

2 works

S. Onstad

2 works

C. Tonielli

3 works

J. L. Green

4 works

C. T. Robinson

2 works

R. Schneider

3 works

K. P. Shen

2 works

Jeremy Keene

2 works

J. H. Langenheim

6 works

B. Fennerty

3 works

Stephanie Zorio

North Carolina Dept of Environmental QualityORCID: 0000-0001-7847-76674 works

A. Kumar

1 work

T. Powell

The University of the South5 works

B. Thomson

2 works

C. G. Curtin

5 works

J. Gutierrez

2 works

Verónica Zepeda

ORCID: 0000-0001-8559-28822 works

B. P. Feifarek

2 works

D. A. Duarte

2 works

Publication (211) →

Show 201 more publications

Quantifying global soil carbon losses in response to warming

2016Naturearticle

On theory in ecology

2014BioSciencearticle

Experimental warming decouples plant-fungal symbiont interactions and leads to a more conservative ecosystem

2026PNASarticle

Plant community composition mediates both large transient decline and predicted long-term recovery of soil carbon under climate warming

2002Global Biogeochemical Cyclesarticle

Global assessment of experimental Climate warming on tundra vegetation: heterogeneity over space and time

2012Ecology Lettersarticle

Plant responses to experimental warming in a montane meadow

2001Ecologyarticle

Integrating experimental and gradient methods in ecological climate change research

2003Ecologyarticle

Microbes on mountainsides: contrasting elevational patterns of bacterial and plant diversity

2008PNASarticle

Biodiversity scales from plots to biomes with a universal species-area curve

2009Ecology Lettersarticle

Climate Warming Drives Local Extinction: Evidence from Observation and Experimentation

2018ScienceAdvancesarticle

Responses of high-altitude graminoids and fungal fungi to 20 years of experimental warming

2014Ecologyarticle

King of the hill? How biotic interactions affect biogeographical pattern and species responses to climate change

2019thesis

Fungal symbionts alter plant responses to global change

2013American Journal of Botanyarticle

Response of plant pathogens and herbivores to a warming experiment

2004Ecologyarticle

Altitudinal gradients fail to predict fungal symbiont responses to warming

2019Ecologyarticle

Effects of long-term experimental warming on aphid density in the field

2007Journal of the Kansas Entomological Societyarticle

The effect of experimental ecosystem warming on CO2 fluxes in a montane meadow

1999Global Change Biologyarticle

Warming disrupts plant–fungal endophyte symbiosis more severely in leaves than roots

2025Global Change Biologyarticle

Reproductive and physiological responses to simulated climate warming for four subalpine species

2007New Phytologistarticle

Shifting macroecological patterns and static theory failure in a stressed alpine plant community

2021Ecospherearticle

Shifts in plant dominance control short and long-term carbon-cycle responses to widespread drought

2006other

The Effect of Climate Change on Soil Organic Carbon over an Elevational Gradient

2022student paper

Compensatory responses to loss of warming-sensitive plant species

2007Ecologyarticle

Propagating climate and vegetation change through the hydrologic cycle in a mountain headwaters catchment.

2015thesis

Changes in ant community composition caused by 20 years of experimental warming vs.13 years of natural climate shift

2014Ecospherearticle

Enhanced growth of sagebrush (<i>Artemisia tridentata</i>) in response to manipulated ecosystem warming

2003Global Climate Changearticle

<i>In situ</i> photosynthetic freezing tolerance for plants exposed to a global warming manipulation in the Rocky Mountains, Colorado, USA

2004New Phytologistarticle

Photosynthetic responses to a climate-warming manipulation for contrasting meadow species in the Rocky Mountains, Colorado, USA

2000Functional Ecologyarticle

Subalpine forest carbon cycling: Short- and long-term influence of climate and species

2005Ecological Applicationsarticle

Herbivory damage but not plant disease under experimental warming is dependent on weather for three subalpine grass species

2023Journal of Animal Ecologyarticle

Context dependent biotic interactions control plant abundance across altitudinal environmental gradients

2019Ecographyarticle

Plant Identity Influences Foliar Fungal Symbionts More Than Elevation in the Colorado Rocky Mountains

2019Microbial Ecologyarticle

Rare species do not disproportionately contribute to phylogenetic diversity in a subalpine plant community

2025American Journal of Botanyarticle

An equation of state unifies diversity, productivity, abundance and biomass

2022Communications Biologyarticle

An Endophyte alters biological characteristics of the grass, Festuca thurberi.

2014student paper

Effects of climate change on growth and seedling establishment of young lodgepole pine

2005student paper

An altitudinal cline in UV floral pattern corresponds with a behavioral change of a generalist pollinator assemblage

2015Ecologyarticle

Traditional plant functional groups explain variation in economic but not size related traits across the tundra biome

2018Global ecology and biogeographyarticle

High-temperature tolerance of <i>Artemisia tridentata</i> and <i>Potentilla gracilis</i> under a climate change manipulation

1996Oecologiaarticle

Experimental examination of early snowmelt-induced water stress in Helianthella quinquenervis: effects on demographic vital rates

2014student paper

Multiple models and experiments underscore large uncertainty in soil carbon dynamics

2018Biogeochemistryarticle

Exploring the impact of climate change on soil carbon storage in montane meadows

2023student paper

Integrating macroecological metrics and community taxonomic structure

2015Ecology Lettersarticle

Pocket gopher (<i>Thomomys talpoides</i>) soil disturbance peaks at mid-elevation and is associated with air temperature, forb cover, and plant diversity

2018Arctic, Antarctic, and Alpine Researcharticle

Control of litter decomposition in a subalpine meadow-sagebrush steppe ecotone under climate change

2001Ecological Applicationsarticle

Maximum information entropy: a foundation for ecological theory

2014Trends in Ecology and Evolutionarticle

Convergence across biomes to a common rain-use efficiency

2004Naturearticle

Soil microbes that may accompany climate warming increase alpine plant production

2019Oecologiaarticle

Consistent shifts in pollinator-relevant floral coloration along a Rocky Mountain elevation gradient

2018Journal of Ecologyarticle

The Changing Floral Color Landscape Across an Alpine Elevation Gradient

2016student paper

Biogeography of plant-associated fungal symbionts in mountain ecosystems: A meta-analysis

2017Diversity and Distributionsarticle

Revisiting Darwin's hypothesis: Does greater intraspecific variability increase species' ecological breadth?

2014American Journal of Botanyarticle

Intraspecific Variation in Responses of a Montane Grass, <i> Festuca thurberi </i>, to Simulated Biological Invasion

2022Frontiers in Forests and Global Changearticle

Sixty-five years of change in montane plant communities in Western Colorado, USA

2016Arctic, Antarctic, and Alpine Researcharticle

Controls on radial growth of mountain big sagebrush and implications for climate change

2009Western North American Naturalistarticle

The effects of Castilleja miniata's parasitic relationship with Delphinium nuttallianum on pollinator visitation and pollination success

2005student paper

Taxon Categories and the Universal Species-Area Relationship

2013The American Naturalist,article

Testing the maximum entropy theory of ecology in the warming meadow

2011student paper

Environmental forces drive morphological variation in an alpine annual plant

2009student paper

Assessing Two Plant Leaf Functional Traits across a Temperature Gradient

2006student paper

Plant dominance in a subalpine montane meadow: biotic vs. abiotic controls of subordinate diversity within and across sites

2018PeerJarticle

Response of nitrogen cycling to simulated climate change: differential responses along a subalpine ecotone

2001Global Change Biologyarticle

The responses of montane shrubs to spatial and temporal climate variation in

2003URBEEarticle

How do above and belowground grass-fungus symbioses change over elevational gradients in mountainous Colorado?

2012student paper

Temperature response of soil respiration largely unaltered with experimental warming

2016Proceedings of the National Academy of Sciencesarticle

Real and experimental ecosystem warming: interacting effects on snowmelt, plant community composition and carbon storage in a Rocky Mountain subalpine meadow

2012student paper

The Direct and Interactive Effects of Warming and Species Interaction on Plant Functional Traits

2019student paper

Testing the predictions of the Maximum Information Entropy Theory for abundance and energy distributions on the Gothic earthflow

2010student paper

Ecosystems impacts of climate change: snowmelt timing, species diversity, and plant productivity

2004student paper

Mountain mosquitoes of the Gothic, Colorado area

1966American Midland Naturalistarticle

Estimating species-area relationships from plot to landscape scale using species spatial-turnover data

1999Oikosarticle

Inferring Regional-Scale Species Diversity from Small-Plot Censuses

2015PLOS ONEarticle

The impact of elevational gradients on dark septate endophytes (DSE) and arbuscular mycorrhizal fungi (AMF) in mountains

2020student paper

Empirical tests of within‐and across‐species energetics in a diverse plant community

2014Ecologyarticle

How do Plant Traits Affect Thermoregulation?

2016student paper

A test of species-area theory on a high disturbance area of the Gothic earthflow

2009student paper

Effects of water addition on above- and below-ground processes in montane meadows

2009student paper

Can fungal symbionts shift host niche dimensions to promote species coexistence?

2011student paper

Devising an ageing technique for <i>Artemisia tridentata ssp. vaseyana</i> near Crested Butte, Colorado

2007student paper

Double keystone bird in a keystone species complex

1993Proceedings of the National Academy of Sciences, USAarticle

Estimating species richness at large spatial scales using data from discrete plots

2004Ecographyarticle

Nest site selection by a secondary cavity-nesting species, the mountain bluebird

2009student paper

Floral pigmentation patterns provide an example of Gloger’s rule in plants

2015Nature Plantsarticle

Intraspecific trait variation affects community distributions of alpine meadow plant communities

2010student paper

The effects of hemiparasitism by <i> Castilleja</i> spp on community structure in alpine ecosystems

2011student paper

Effects of experimental warming on herbivory and pathogen loads on subalpine grass species Festuca thurberi, Poa pratensis, and Achnatherum lettermanii

2015student paper

Aboveground and belowground traits of dominant and sub-dominant species along an elevational gradient.

2014student paper

The Effects of Ants on Ecosystem Dynamics: Investigating the Ecological Influence of Ant Nests in the Rocky Mountains

2016student paper

The impacts of environmental change on plant and microbial communities: A turf transplant experiment in the Colorado Rocky Mountains

2025thesis

Pocket gopher activity across elevation gradients.

2014student paper

Mix and Match: Transplanting symbiotic fungal partners across elevational gradients to gauge responses in migrating Elymus hosts

2018student paper

Plant Successional Changes Over 67 Years on the Gothic Earthflow.

2014student paper

Identifying the Impact of Biotic Interactions on Meadow Species Distributions in Sagebrush and Bunchgrass Dominated Systems

2019student paper

Plant succession on a subalpine earthflow in Colorado

1956Ecologyarticle

Willow branch selection by the red-naped sapsucker (<i>Sphyrapicus nuchalis</i>): The effects of branch characteristics on foraging behavior in Gothic, CO

2010student paper

Range-wide study in a sexually polymorphic wild strawberry reveals climatic and soil associations of sex ratio, sexual dimorphism and sex chromosomes

2025Journal of Ecologyarticle

The effect of climate change on the germination and growth rates of young subalpine fir (<i>Abies lasiocarpa</i>)

2004student paper

Does nutrient enrichment interact with <i>Castilleja miniata</i> to alter plant communities in montane meadows?

2012student paper

The effects of plant succession and ant nest on soil organic matter and soil moisture

2002student paper

An Epichloe endophyte alters the ecology of Poa leptocoma.

2014student paper

Getting to the Root of It: Effects of Castilleja Root Hemiparasitism on Plant Community Structure and Function

2024student paper

Red-naped Sapsucker nest tree selection and effects on the willow breeding bird community

2004student paper

Factors influencing floral traits in Rocky Mountain meadows: competition, environmental filtering, and phylogeny

2009student paper

Tracing phenology trends in time and space using herbarium data and citizen science observations in western Colorado

2020student paper

Can fungal symbionts shift host niche dimensions to promote species coexistence?

2013student paper

Effects of proximity to riparian zones on avian species richness and abundance in montane aspen woodlands.

2015Journal of Field Ornithologyarticle

Metabolic Partitioning across Individuals in Ecological Communities

2017Global Ecology and Biogeographyarticle

Defensive strategy to nectar robbing in <i></i>Ipomopsis aggregata:<i></i> floral nectar as a tolerance trait

2003student paper

The effect of sampling effort on species richness estimates of flower visitors

2007student paper

Intraspecific variation of specific leaf area along an elevational gradient

2011student paper

A Brief Escape to Normalcy: A Summer at RMBL

2020Mountain Views Chronicleother

The response of four subalpine forbs to supplemental nitrogen within different soil moisture environments

2009student paper

The future of plant-fungal symbioses along elevational gradients

2016student paper

Attributes of mountain bluebird cavity nests

2009student paper

Competitive hierarchy or niche packing? An examination of the effects of competition on community.

2014student paper

Plant community composition with respect to species richness along an altitudinal gradient in gunnison county, co

2009student paper

Species richness along an elevational gradient

2009student paper

The impacts of long term warming on potential soil microbial activity across soil depth

2019student paper

How do distributions of belowground grass-fungal symbioses change over altitudinal gradients in the Colorado Rocky Mountains?

2015student paper

Tracking diverse minerals, hungry organisms, and dangerous contaminants using reactive transport models

2019Elementsarticle

Surveying historical patterns in vegetation change (1929-2019) in the upper East River Valley using RMBL's archival herbarium records

2019student paper

Global warming and terrestrial ecosystems: a conceptual framework for analysis

2000BioSciencearticle

A Comparative Analysis of Saxicolous Lichen Diversity and Cover along an Elevation Gradient on Talus Slopes in Gunnison National Forest to Infer Possible Climate Change Effects

2017student paper

Climate change and extinction risk

2004Naturearticle

Variation in root morphology and allocation of <i>Androsace septentrionalis</i> along a natural elevation/climate gradient

2013student paper

Patterns of biodiversity in sub-alpine wetlands

2006student paper

The effects of climate change on subalpine fir (<i></i>Abies lasiocarpa<i></i>) sapling growth and establishment success across an elevational gradient

2006student paper

Insight from Integration

2012Naturearticle

Phenology of Arbuscular Mycorrhizal fungi and Dark Septate Endophytes across an elevation gradient.

2014student paper

Behavioral response of Mountain White-Crowned Sparrows towards an interspecific competitor

2013student paper

Effects of water addition on below- and above-ground carbon processes across a montane elevational gradient

2008student paper

Intraspecific flower color variation along elevation gradients

2019student paper

How does experimental warming effect the rate of herbivory and fungi on host grasses?

2017student paper

Intraspecific signaling function of Crown Coloration in Mountain White-Crowned Sparrows

2005student paper

What sexual diversity in the natural world can teach humans about sex, gender, and inclusive biology

2024BioSciencearticle

Plant size influences mycorrhizal colonization of <i>Polemonium foliosissimum</i>

2011student paper

Scale collapse and the emergence of the power law species–area relationship

2015Global Ecology and Biogeographyarticle

Changes in water relations for leaves exposed to a climate-warming manipulation in the Rocky Mountains of Colorado

1997Environmental and Experimental Biologyarticle

Impacts of the invasive species <i> Linaria vulgaris </i> on soil ecology and native plant biodiversity

2019student paper

Dead wood biomass and turnover time, measured by radiocarbon, along a subalpine elevation gradient

2004Oecologiaarticle

Responses of subalpine meadow vegetation to four years of experimental warming

2000Ecological Applicationsarticle

Dormancy rates in microbial communities across an elevational gradient

2017student paper

Dispersal, phenology and habitat preference of subalpine whirligig beetles (Coleoptera: Gyrinidae: Gyrinus)

2010student paper

On the spatial distribution and abundance of herbaceous plants in aspen and conifer forest understories: a test of neutral/statistical versus niche-based theories?

2008student paper

How do riparian songbirds (Passerines) coexist? Effects of vegetative structural complexity on habitat selection in willow (<i>Salix</i> spp.)

2012student paper

The Gothic earthflow revisited: a chronosequence examination of colonization on a subalpine earthflow

1994Vegetatioarticle

Influence of patch area on bird species diversity in coniferous forests

2009student paper

Climate change and anti-herbivory resistance communication in <i>Artemisia tridentata</i>

2007student paper

Understanding Organismal Capacity to Respond to Anthropogenic Change: Barriers and Solutions

2021Integrative and Comparative Biologyarticle

A mutualistic endophyte alters the niche dimensions of its host plant

2015AoB Plantsarticle

Moss and vascular plant cover across elevational gradients in a changing alpine climate

2025student paper

Talus turnover: A study of the distribution of lichens along elevational gradients

2013student paper

On the implications of species-area relationships for endemism, spatial turnover, and food web patterns

1997Oikosarticle

Won't you be my neighbor: neighborhood effect influences mycorrhizal and endophyte colonization

2013student paper

Legacy effects and arbuscular mycorrhizal fungi of Linaria vulgaris invasion in Colorado and Illinois, USA

2025Invasive Plant Science and Managementarticle

Gas exhange and water relations of two Rocky Mountain shrub species exposed to a climate change manipulation

2000Plant Ecologyarticle

Trap behavior of <i> Zonotrichia leucophrys oriantha </i> at different elevations in the West Elk Mountains, Colorado

2019student paper

Factors that affect territory size in Mountain White-Crowned Sparrows (<i>Zonotrichia leucophrys oriantha</i>)

2017student paper

Changes in mosquito community of Gunnison County since 1966

2007student paper

Ant presences alter plants: The impact <i>Formica obscuripes</i> presence on plants and soil micro-anthropod diversity

2011student paper

Habitat patch use, density, and territoriality of American Red Squirrels (<i>Tamiasciurus hudsonicus</i>) in the southwestern Rocky Mountains, Colorado

2016student paper

Environmental Variation and Vegetative Composition in Beaver Engineered Ecosystems

2024student paper

Determining Long-Term Success of Revegetation Efforts in Disturbed Sites

2009student paper

What determines the distribution of red-naped sapsuckers in the East River Valley?

2008student paper

Revegetation after disturbance in high-altitude meadow ecosystems

2006student paper

Biotic and abiotic effects on ant community structure

2002student paper

Dichotomous key to the members of the onagraceae family found in the gothic area

2009student paper

the influence of red-naped sapsuckers on willow bird communities

2005student paper

Experimental test of the ability of plants to regulate their temperatures under climate change: Using transplanted communities, thermal imaging, and arcGIS across an elevation gradient

2021student paper

Investigating the air temperatures of different taxa in an alpine plant community

2021student paper

Predictors and Strength of Microclimate Buffering in the Gunnison Valley

2020student paper

The effects of elevation and climate on butterfly (<i>Lepidoptera</i>) abundance and activity

2011student paper

Understanding how leaf endophytes are affected by climate change: Examining fungi in grass species with warming

2016student paper

Effects of Altitude on the Growth, Mortality, and Recruitment of Subalpine Tree Species in the West Elk Mountains of Colorado.

2023student paper

Investigating the comparison between bird diversity and apex aquatic predators in sub-alpine beaver ponds

2021student paper

Host effects on herbivory and pollination in a hemiparasitic plant

2002Ecologyarticle

Carbon Cycle Uncertainty Increases Climate Change Risks and Mitigation Challenges

2012Journal of Climatearticle

Possible influence of altitude, geographical distance between sites and annual precipitation rates on species richness

2009student paper

Aspen heart rot fungus (<i>Phellinus tremulae</i>) distribution in aspen forests in relation to open meadows: implications for red-naped sapsucker (<i>Sphyrapicus nuchalis</i>) nesting habitat

2008student paper

Building up Biogeography: Pattern to Process

2018Journal of Biogeographyarticle

Effect of Rock Size, Age, and Distance on the Biodiversity of Lower Copper Creek Area Lichens: A Test of the Theory of Island Biogeography

2004student paper

Land Use and Landscape Change in the Colorado Mountains II: A Case Study of the East River Valley

1996Mountain Research and Developmentarticle

Host specificity of hemiparasitic Castilleja and its influence on plant community diversity.

2023student paper

Effects of Alpine Shrub Cover on Territorial Male White-Crown Sparrows Densities.

2015student paper

Does a foliar endophyte improve plant fitness under flooding?

2017Plant Ecologyarticle

Species coexistence in a food web with intraguild predation

2004URBEEarticle

Avian Diversity in Aspen and Conifer Forests :

2004student paper

Untangling multiple factors in spatial distributions: Lilies, gophers, and rocks

1996Ecologyarticle

Comparison of Arthropod Diversity between Perennial Shrub versus Evergreen Tree Habitats at Sites Near Gothic, CO

2004student paper

RELATIONSHIP BETWEEN ASPEN HEARTWOOD ROT AND THE LOCATION OF CAVITY EXCAVATION BY A PRIMARY CAVITY-NESTER, THE RED-NAPED SAPSUCKER

2006The Condorarticle

Effects of manipulated soil microclimate on mesofaunal biomass and diversity

1996Soil Biology and Biochemistryarticle

Population dynamics and competitive outcome derive from resource allocation statistics: the governing influence of the distinguishability of individuals

2015Theoretical Population Biologyarticle

The shape of a species\\\' spatial abundance distribution

2012Global Ecology and Biogeographyarticle

Acidification and salamander recruitment

1994BioSciencearticle

Non-random orientation of entrance holes to woodpecker nests in aspen trees

1976Condorarticle

The responses of lake waters to organic matter additions.

1983Hydrobiologiaarticle

Toward a synthesis of the Newtonian and Darwinian worldviews

2002Physics Todayarticle

Visitors to Red-Naped Sapsucker sap wells and the effectiveness of artificial wells

2025student paper

Habitat of Pocket Gophers in Cochetopa Creek Drainage, Colorado

1968American Midland Naturalistarticle

Moisture Relationships in Twelve Northern Desert Shrub Communities Near Grand Junction, Colorado

1976Ecologyarticle

Can montane landscapes recover from human disturbance? Long-term evidence from disturbed subalpine communities

1995Biological Conservationarticle

Dataset (26) →

NEON AOP Survey of Upper East River CO Watersheds: Waveform LiDAR Binary Data

The waveform Light Detection and Ranging (LiDAR) data in this package were generated through a National Ecological Observatory Network Airborne Observ...

ess_dive2024

Range size and local abundance data for angiosperm communities across an elevation gradient, Rocky Mountain Biological Lab, 2021-2022

This dataset contains abundance and range size data for angiosperm communities at three sites in Washington Gulch near the Rocky Mountain Biological L...

other2025

Data from: 'Abiotic influences on continuous conifer forest structure across a subalpine watershed'

This package archives the core data used for analysis and inference in 'Abiotic influences on continuous conifer forest structure across a subalpine w...

ess_dive2025

Data for Context-dependent biotic interactions control plant abundance across altitudinal environmental gradients, 2014, 2016, Colorado, USA

Many biotic interactions influence community structure, yet most distribution models for plants have focused on plant competition or used only abiotic...

other2020

Langenheim Plant Species Data (1953) and Associated Resurvey Datasets (2014), Gunnison Basin, Colorado, USA

Quantitative plant abundance data were collected from the same 121 sites at two time periods separated by 65 years (1948-1952 and 2012-2014) in the Co...

other2025

Data for 'Weak latitudinal gradients in insect herbivory for dominant rangeland grasses of North America'

Patterns of insect herbivory may follow predictable geographical gradients, with greater herbivory at low latitudes. However, biogeographic studies of...

other2020

Data for Context-dependent biotic interactions control plant abundance across altitudinal environmental gradients, 2014, 2016, Colorado, USA.

Many biotic interactions influence community structure, yet most distribution models for plants have focused on plant competition or used only abiotic...

other2021

Mammalian herbivores restrict the altitudinal range limits of three alpine grass species

Though rarely experimentally tested, biotic interactions have long been hypothesized to limit low-elevation range boundaries of species. We tested t...

other2023

Data for Lynn et al. “Soil microbes that may accompany climate warming increase alpine plant production”

Climate change is causing species with non-overlapping ranges to come in contact, and a key challenge is to predict the consequences of such species r...

other2020

An Equation of State Unifies Diversity, Productivity, Abundance and Biomass (data sets)

[object Object]

2022
Show 16 more datasets

Plant composition data from 67 grassland sites of the Upper Gunnison Basin, CO, USA, 2014

Here, we deposit data from a vegetation survey conducted in 2014. The data was collected to document current vegetation patterns in the region, parame...

other2020

Langenheim Plant Species Data (1953) and Associated Resurvey Datasets (2014), Gunnison Basin, Colorado, USA

Quantitative plant abundance data were collected from the same 121 sites at two time periods separated by 65 years (1948-1952 and 2012-2014) in the Co...

other2025

Annually collected demography data from an alpine plant community on Mt. Baldy, Colorado (38.978725°N, 107.042104°W, ~3540 masl).

Description: Annual demography dataset for an alpine plant community in Colorado. This file updates previous years of data for this project posted to ...

other2021

Appendix A. List of the graminoid species in the experimental warming meadow, Rocky Mountain Biological Laboratory, Gunnison County, Colorado, USA.

List of the graminoid species in the experimental warming meadow, Rocky Mountain Biological Laboratory, Gunnison County, Colorado, USA.

other2016

Pocket gopher (<i>Thomomys talpoides</i>) soil disturbance peaks at mid-elevation and is associated with air temperature, forb cover, and plant diversity

Burrowing mammals can be ecosystem engineers by increasing soil aeration and erosion and altering the structure of plant communities. Studies that cha...

other2019

Data for “Herbivory damage but not plant disease under experimental warming is dependent on weather for three subalpine grass species”, Rocky Mountain Biological Laboratory, Gothic, Colorado, 2015-2017.

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2022

1 m Resolution Multiscale Height-above-stream Wetness Index for the Upper Gunnison Domain

<p>This map is a soil moisture proxy derived from analysis of the UG 1m hydrologically corrected digital elevation model. The intuition behind this ma...

s32021

Biomass Inventories at Harvard Forest EMS Tower since 1993

In 1993, we installed 40 circular, 10 m radius biometric plots in the footprint of the EMS tower on Prospect Hill. We randomly placed the plots within...

other2021

Forage Resources in Warming and Removal Plots, Almont, CO, 2019

This is data collected to explore the impacts of warming and dominant species removal on the quantity and quality of plants for cattle foraging. The d...

other2026

Forage Resources in Warming and Removal Plots, Almont, CO, 2019

This is data collected to explore the impacts of warming and dominant species removal on the quantity and quality of plants for cattle foraging. The d...

other2026

Supplemental Tables for "Phylogenetic patterns over sixty-five years of vegetation change across a montane elevation gradient" by Veldhuisen et al.

This repository contains three supplemental tables for Veldhuisen et al.'s paper titled "Phylogenetic patterns over sixty-five years of vegetation cha...

other2025

Model output and meta-analysis data from INTERFACE paper

Model output and meta-analysis data from model-experiment comparison that came out of INTERFACE workshop. Includes output from five soil carbon models...

other2018

Forest Inventory of a Northern Hardwood Forest: Watershed 6, 2017, Hubbard Brook Experimental Forest

Forest inventory surveys were initiated in 1965, repeated in 1977, and repeated at 5 year intervals after that; this data set was collected in summer ...

other2019

RMBL Digitized Herbarium Records

Vascular Plants curated within the Rocky Mountain Biological Laboratory Herbarium. Specimens date to 1929 and are primarily from the East River and up...

other2020

Data from: Medical-device recalls in the UK and the device-regulation process: retrospective review of safety notices and alerts

Background: Medical devices are used widely for virtually every disease and condition. Although devices are subject to regulation, the number of recal...

other2011

Appendix A. List of the graminoid species in the experimental warming meadow, Rocky Mountain Biological Laboratory, Gunnison County, Colorado, USA.

List of the graminoid species in the experimental warming meadow, Rocky Mountain Biological Laboratory, Gunnison County, Colorado, USA.

other2016